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The Garden strawberry (Fragaria
×ananassa and related
cultivars) is the most
common variety of strawberry cultivated worldwide. Like
other species of
Fragaria (strawberries),
it belongs to the family
Rosaceae;
its fruit is more technically known as an
accessory fruit, in that the
fleshy part is derived not from the plant's ovaries (achenes)
but from the peg at the bottom of the bowl-shaped
hypanthium
that holds the ovaries.
The
Garden Strawberry was first bred in Europe in the early
18th century, and represents the accidental cross of
Fragaria virginiana
from eastern North America,
which was noted for its fine flavor, and
Fragaria chiloensis
from Chile
and noted for its large size.
Cultivars of Fragaria ×ananassa have replaced in
commercial production the
Woodland Strawberry, which was the first strawberry
species cultivated in the early 17th century.
History
The
typical cultivated strawberry comes from the
Americas,
and is a hybrid of the North America F. virginiana
and the South American F. chiloensis, developed
in the early 18th century. Interestingly, the
crossbreeding was done in Europe to correct a mistake;
the European
horticulturists had only
brought female South American plants, and were forced to
cross them with the North American variety in order to
get fruit and seeds. The name Fragaria comes from "fragans",
meaning odorous, referring to the perfumed flesh of the
fruit. Madam Tallien,
a great figure of the
French Revolution, who was nicknamed Our Lady of
Thermidor,
used to take baths full of strawberries to keep the full
radiance of her skin.
Fontenelle, centenarian
writer and gourmet of the 18th century, considered his
long life was due to the strawberries he used to eat.
Though originally from the New World, strawberries were
considered poisonous in Argentina until the mid-nineteenth
century.
Popular etymology
has it that the name "straw" berry comes from gardeners'
practice of mulching
strawberries with straw
to protect the fruits from rot (a pseudoetymology that
can be found in non-linguistic sources such as the
Old Farmer's Almanac 2005).
There is
an alternative, albeit equally implausible, theory that
the name derives from the Anglo-Saxon verb for "strew" (meaning
to spread around) which was streabergen (Strea means "strew"
and Bergen means "berry" or "fruit") and thence to
streberie, straiberie, strauberie, straubery, strauberry,
and finally, "strawberry", the word which we use today.
The name might have come from the fact that the fruit
and various runners appear "strewn" along the ground.
However, there is no evidence that the Anglo-Saxons ever
grew strawberries, and even less that they knew of this
practice, as strawberries are originally from the
America
Fragaria x ananassa
'Gariguette', a cultivar grown in southern
France.
Strawberry varieties vary remarkably in size, color,
flavor, shape, degree of fertility, season of ripening,
liability to disease and constitution of plant. Some
vary in foliage, and some vary materially in the
relative development of their sexual organs. In most
cases the flowers appear hermaphroditic in structure,
but function as either male or female.
For
purposes of commercial production, plants are propagated
from runners and generally distributed as either bare
root plants or plugs. Cultivation follows one of two
models, annual plasticulture or a perennial system of
matted rows or mounds. A small amount of strawberries
are also produced in greenhouses during the off season.
The bulk
of modern commercial production uses the plasticulture
system. In this method, raised beds are formed each year
and covered with black plastic, which prevents weed
growth and erosion, under which is run irrigation tubing.
Plants, usually obtained from northern nurseries, are
planted through holes punched in this covering. Runners
are removed from the plants as they appear, to encourage
the plants to put most of their energy into fruit
development. At the end of the harvest season, the
plastic is removed and the plants are plowed into the
ground. Because strawberry plants more than a year or
two old begin to decline in productivity and fruit
quality, this system of replacing the plants each year
allows for improved yields and more dense plantings.
However, because it requires a longer growing season to
allow for establishment of the plants each year, and
because of the increased costs in terms of forming and
covering the mounds and purchasing plants each year, it
is not always practical in all areas.
Ripe fruit of the Garden Strawberry
The
other major method is to maintain the same plants from
year to year. The runners of established plants should
be allowed to root in the soil adjoining the plants,
which should, therefore, be kept light and fine, or
layered into small pots as for forcing. As soon as a few
leaves are produced on each the secondary runners should
be stopped. When the plants have become well-rooted they
should at once be planted out. They do best in a rather
strong loam,
and should be kept tolerably moist. The ground should be
trenched 50-100 cm deep, and supplied with plenty of
manure, a good proportion of which should lie just below
the roots, 25-30 cm from the surface. The plants may be
put in on an average about 50-60 cm apart.
The
plantation should be renewed every second or third year,
or less frequently if kept free of runners, if the old
leaves are cut away after the fruit has been gathered,
and if a good top-dressing of rotten dung or leaf mold
is applied. A top-dressing of loam is beneficial if
applied before the plants begin to grow in spring, but
after that period they should not be disturbed during
the summer either at root or at top. If the plants
produce a large number of flower-scapes, each should, if
fine large fruit is desired, have them reduced to about
four of the strongest. The lowest blossoms on the scape
will be found to produce the largest, earliest and best
fruits. The fruit should not be gathered until it is
quite ripe, and then, if possible, it should be quite
dry, but not heated by the sun. Those intended for
preserving are best taken without the stalk and the
calyx.
A
mulching of straw manure
put between the rows in spring serves to keep the ground
moist and the fruit clean, as well as to afford
nourishment to the plants. Unless required, the runners
are cut off early, in order to promote the swelling of
the fruit. The plants are watered during dry weather
after the fruit is set, and occasionally until it begins
to colour. As soon as the fruit season is over, the
runners are again removed, and the ground hoed and raked.
Fragaria × ananassa
'Chandler' , a short day commercial variety
grown in
California.
Strawberries are often grouped according to their
flowering habit. Traditionally, this has consisted of a
division between "June-bearing" strawberries, which bear
their fruit in the early summer and "Ever-bearing"
strawberries, which often bear several crops of fruit
throughout the season. More recently, research has shown
that strawberries actually occur in three basic
flowering habits: short day, long day, and day neutral.
These refer to the day length sensitivity of the plant
and the type of photoperiod which induces flower
formation. Day neutral cultivars produce flowers
regardless of the photoperiod. Most commercial
strawberries are either short day or day neutral.
While
rarely if ever done commercially, strawberries may also
be propagated by seed, and a few seed propagated
cultivars have been developed for home use. Seeds are
acquired commercially or saved from fruit ripened early
in the summer. They may at once be sown, either in a
sheltered border outdoors or in pots, or better in March
under glass, when they will produce fruits in June of
the same year. The soil should be rich and light, and
the seeds very slightly covered by sifting over them
some leaf-mould or old decomposed cow
dung.
When the plants appear and have made five or six leaves,
they are transplanted to where they are to remain for
bearing. The seeds sown in pots may be helped on by
gentle heat, and when the plants are large enough they
are pricked out in fine rich soil, and in June
transferred to the open ground for bearing.
Pollination
Most cultivars are somewhat self fertile, but good
bee
activity has been shown to improve
pollination,
which results in larger and better shaped berries.
Commercial growers sometimes place
beehives
within range of the fields to increase bee populations.
Forcing
The
runners propagated for forcing are layered into 75 mm
pots, filled with rich soil, and held firm by a piece of
raffia,
a peg or stone. If kept duly watered they will soon form
independent plants. The earlier they are secured the
better.
When
firmly rooted they are removed and transferred into well-drained
150 mm pots, of strong well-enriched loam, the soil
being rammed firmly into the pots, which are to be set
in an open airy place. In severe frosts they should be
covered with dry litter or bracken, but do not
necessarily require to be placed under glass. They are
moved into the forcing houses as required.
The main
points to be kept in view in forcing strawberries are:
- use
strong stocky plants, the leaves of which have grown
sturdily from being well exposed to light, and
- grow
them slowly until the fruit is set.
1890
watercolor
of strawberries (cultivar 'Parker Earle').
When they
are first introduced into heat, the temperature should
not exceed 8°C to 10°C, and air must be freely admitted;
should the leaves appear to grow up thin and delicate,
less fire heat and more air must be given, but an
average temperature of 13°C by day may be allowed and
continued while the plants are in flower.
When the
fruit is set the heat may be gradually increased, till
at the ripening period it stands at 18°C to 24°C by sun
heat. While the fruit is swelling the plants should
never be allowed to get dry, but when it begins to
colour no more water should be given than is absolutely
requisite to keep the leaves from flagging. The plants
should be removed from the house as soon as the crop is
gathered. The forced plants properly hardened make first-rate
outdoor plantations, and if put out early in summer, in
good ground, will often produce a useful autumnal crop.
Pests
A
number of species of
Lepidoptera feed on
strawberry plants; for details see
this list.
Diseases
The
most troublesome fungoid attacks to which the strawberry
is subject are mildew,
leaf spot and
leaf blight. The former, like all
mildews, attacks the leaves and spreads to the fruit,
these being covered with the white mycelium. The fungus
is identical with that causing mildew in
hops
(Sphaerotheca humuli), and its development is
greatly furthered by exposure of its host to cold
draughts or low night temperatures. Spraying the foliage
with potassium
sulfide (K2S) (mixed with water at a 1:40
ratio by volume) should hold it in check, but the plants
should not be sprayed when the fruit is developing.
Leaf
spot is caused by the fungus
Sphaerella fragariae.
The first symptom of this attack is the appearance of
small, circular, white spots on the leaves, having a
broad, definite, dark reddish margin.
On
these spots a whitish mould develops, and this is
followed later by the perfect form of the fungus, the
fruits of which appear to the naked eye as small black
spots seated on the white dead spot on the leaf.
Potassium sulfide may be used as for the mildew, or,
perhaps better, Bordeaux
mixture. Some recommend cutting off the leaves after
fruiting and turn the beds over so as to destroy the
fungus in the leaves.
Leaf
blight is caused by the fungus
Phomopsis obscurans.
The symptoms begin as one to several circular reddish-purple
spots on a leaflet. Spots enlarge to V-shaped lesions
with a light brown inner zone and dark brown outer zone.
Lesions follow major veins progressing inward. A copper
sulphate containing spray like
Bordeaux mixture will help control
this disease.
The
grubs
of the cockchafer
(Meloloniha vulgaris) and the rose chafer (Cetonia
aurata) frequently feed
upon the roots of the strawberry and do considerable
damage, while the larvae
of the Ghost Moth (Hepialus
humuli) and garden swift moth behave in a similar
way. The imago
of Cetonia aurala also frequently damages the
flowers of the strawberry by devouring their centres,
and is often troublesome in this way in forcing-houses
particularly. The carnivorous
ground beetles, particularly
Pterostichus nigra and Harpalus rufimanus,
when the fruit is ripe attack it at night, returning to
the soil in the daytime. They are to be caught by
placing jars containing some attractive matter, such as
meat and water, at intervals about the beds with their
mouths sunk level with the surface of the soil.
Millipedes
also are often found in the ripe fruit, but occur mostly
where the soil is very rich in organic matter and poor
in lime. Slugs
and snails
also snack on the fruit, as do
birds.
Slug pellets can be used to reduce their numbers, with
child and animal safe versions available. Organic
solutions to slug attacks include beer baiting.
In addition to
being consumed fresh, strawberries are frozen or made
into preserves.
Strawberries are a popular addition to dairy products,
as in strawberry flavored ice cream, milkshakes and
yogurts. Strawberry pie is also popular. Strawberries
can also be used as a natural acid/base indicator. They
are also supposedly used for whitening teeth.
Nutrition
One
cup (144 g) of strawberries constitutes approximately 45
calories (188 kJ) and is an excellent source of vitamin
C and flavonoids.
|
Nutrient |
Units |
1 cup, whole
-------
144 g |
|
Proximates |
|
Water |
g |
132 |
|
Energy |
kcal |
43 |
|
Energy |
kJ |
181 |
|
Protein |
g |
0.88 |
|
Total lipid (fat) |
g |
0.53 |
|
Carbohydrate, by difference |
g |
10.1 |
|
Fiber, total dietary |
g |
3.3 |
|
Ash |
g |
0.62 |
|
Minerals |
|
Calcium, Ca |
mg |
20 |
|
Iron, Fe |
mg |
0.55 |
|
Magnesium, Mg |
mg |
14 |
|
Phosphorus, P |
mg |
27 |
|
Potassium, K |
mg |
240 |
|
Sodium, Na |
mg |
1.44 |
|
Zinc, Zn |
mg |
0.19 |
|
Copper, Cu |
mg |
0.07 |
|
Manganese, Mn |
mg |
0.42 |
|
Selenium, Se |
µg |
1.01 |
|
Vitamins |
|
Vitamin C, ascorbic acid |
mg |
82 |
|
Thiamin |
mg |
0.03 |
|
Riboflavin |
mg |
0.10 |
|
Niacin |
mg |
0.33 |
|
Pantothenic acid |
mg |
0.49 |
|
Vitamin B-6 |
mg |
0.09 |
|
Folate |
µg |
25 |
|
Vitamin B-12 |
µg |
0 |
|
Vitamin A, IU |
IU |
39 |
|
Vitamin A, RE |
µg RE |
4.3 |
|
Vitamin E |
mg ATE |
0.20 |
|
Nutrient |
Units |
1 cup, whole
-------
144 g |
|
Lipids |
|
Fatty acids, saturated |
g |
0.03 |
|
16:0 |
g |
0.02 |
|
18:0 |
g |
0.006 |
|
Fatty acids, monounsaturated |
g |
0.075 |
|
16:1 |
g |
0.001 |
|
18:1 |
g |
0.073 |
|
Fatty acids, polyunsaturated |
g |
0.27 |
|
18:2 |
g |
0.16 |
|
18:3 |
g |
0.11 |
|
Cholesterol |
mg |
0 |
|
Phytosterols |
mg |
17 |
|
Amino acids |
|
Tryptophan |
g |
0.01 |
|
Threonine |
g |
0.027 |
|
Isoleucine |
g |
0.02 |
|
Leucine |
g |
0.045 |
|
Lysine |
g |
0.036 |
|
Methionine |
g |
0.001 |
|
Cystine |
g |
0.007 |
|
Phenylalanine |
g |
0.026 |
|
Tyrosine |
g |
0.030 |
|
Valine |
g |
0.026 |
|
Arginine |
g |
0.037 |
|
Histidine |
g |
0.017 |
|
Alanine |
g |
0.045 |
|
Aspartic acid |
g |
0.20 |
|
Glutamic acid |
g |
0.13 |
|
Glycine |
g |
0.035 |
|
Proline |
g |
0.027 |
|
Serine |
g |
0.033 |
|